They appear in the following combats:
- The Pyrewings
- The Penitent
- Conduit Masters
- The Shadewings
- Seraph the Temperant
- Seraph the Chaste
- Seraph the Diligent
- Seraph the Patient
- (DLC) The Last Divinity: Divine Intervention
Their focus on defense has caused them to evolve into smaller beings with oversized wings, making them perfect shields for the rest of Seraph's army. We've found brute force is the best way to get past them.
- A Steelwings always appears on Ring 8, at the 2nd to last wave.